- BS - With High Distinction (1971) Pennsylvania State University
- MD Jefferson Medical College, Thomas Jefferson University
- Internship - Internal Medicine Thomas Jefferson University
- Resident - Internal Medicine Mayo Graduate School of Medicine, Mayo Clinic College of Medicine
- Fellow - Gastroenterology Mayo Graduate School of Medicine, Mayo Clinic College of Medicine
- Research Training - NIH Research Trainee, Gastroenterology Unit Mayo Graduate School of Medicine, Mayo Clinic College of Medicine
- MS - Medicine University of Minnesota
- Fellow - Post-doctoral Fellow/Post-doctoral Research Training in Cell Biology Yale University School of Medicine
- Professor of Biochemistry/Molecular Biology
- Professor of Medicine
- Pancreatic disease
- Islet tumors
- Zollinger-Ellison syndrome
See a listing of publications on PubMed, a service of the National Library of Medicine.
See a description of research activities.