- BS - Chemistry-Physiology University of Illinois at Urbana-Champaign
- MD - Medicine University of Illinois, Peoria
- Resident - Internal Medicine Mayo Graduate School of Medicine, Mayo Clinic College of Medicine
- Resident - Thoracic Disease Mayo Graduate School of Medicine, Mayo Clinic College of Medicine
- Resident - Research Fellowship Washington University
- Associate Professor of Biochemistry/Molecular Biology
- Professor of Medicine
See a listing of publications on PubMed, a service of the National Library of Medicine.
See a description of research activities.